http://www.ncbi.nlm.nih.gov/pubmed/11260506
HLA genes in Macedonians and the sub-Saharan origin of the Greeks.
Arnaiz-Villena A, Dimitroski K, Pacho A, Moscoso J, Gómez-Casado E, Silvera-Redondo C, Varela P, Blagoevska M, Zdravkovska V, Martínez-Laso J.
Greeks are found to have a substantial relatedness to sub-Saharan (Ethiopian) people, which separate them from other Mediterranean groups. Both Greeks and Ethiopians share quasi-specific DRB1 alleles, such as *0305, *0307, *0411, *0413, *0416, *0417, *0420, *1110, *1112, *1304 and *1310. Genetic distances are closer between Greeks and Ethiopian/sub-Saharan groups than to any other Mediterranean group and finally Greeks cluster with Ethiopians/sub-Saharans in both neighbour joining dendrograms and correspondence analyses.

